An rat named Remy dreams to become a French boxer despite his family's wishes and the problem to be a rat at a profession. He discovers himself situated beneath a restaurant made famous by his hero, Auguste Gusteau when fate puts Remy in the sewers of Paris. Inspite of the dangers of as an improbable - and unwanted - tourist at a nice restaurant's kitchen, Remy's passion for cooking sets a pit of debt into the world of Paris turns down.